I don´t agree with that, okay, to a certain extent if you focus on body design, pickup layout. To hunt in Strat´s territory it is face validity to have a pickup layout with a singlecoil in the middle between a bass and a treble pickup. I am aswell not convinced that this is very much related to material or neck construction (bold on or set neck), but scale is a factor.
The same with Tele.
I tested a bunch of Teles (MiUS, MiM, MiJ, Squier (China, other Asian countries)) with SC SC layout and wasn´t convinced by that supposed unique singlecoil sound.
And it´s the same with all those LP enthusiasts seeking the holy grail sound...
I found a guitar which gives me warm (and if necessary fat) humbucker sound for all kind of clean, crunch, overdrive, high gain. But it deals with snappy singlecoil voicings for dirty crunch or even clean sounds, dealing with familiar voicings for position 2 and 4.
Recently I purchased an other guitar designed by PRS and built in Korea, with only has a HB HB layout, but coil split action via push-pull puti. And in singlecoil mode this guitar is very nice an snappy, too. And it´s not a Tele or Strat. It has a quarter more scale than my Core one. And it´s exactly with that in Fender territory.
My two Cent.